Fossil footprints discovered in southeastern Poland are 395 million years old, making them the oldest known tetrapod tracks and the oldest evidence of true tetrapods.
Sources
- Oldest Land-Walker Tracks Found--Pushes Back Evolution (nationalgeographic.com)
- https://www.cnn.com/2010/WORLD/europe/01/07/tetrapods.poland.evolution.discovery/index.html (cnn.com)
- https://www.nature.com/articles/nature08623 (nature.com)
- https://www.science.org/content/article/ancient-four-legged-beasts-leave-their-mark (science.org)
- https://www.earthmagazine.org/article/tetrapod-tracks-reset-timing-four-legged-evolution/ (earthmagazine.org)
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